Following a cataclysmic conflict known as the Sixty Minute War, the remnants of humanity regroup and form mobile "predator" cities. Under a philosophy known as "Municipal Darwinism", larger cities hunt and absorb smaller settlements in the "Great Hunting Ground", which includes Great Britain and Continental Europe. In opposition, an "Anti-Traction League" have developed an alternative civilization consisting of "static settlements" in Asia led by Shan Guo (formerly China), protected by the "Shield Wall." A young girl named Hester Shaw works alongside a young historian to take down London, once and for all.
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